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News & Upcoming Events


Harvard study finds that nice guys do finish first!

PED is newly WED
(or WEDDED?)

New York Times article on Martin Nowak

Boston Globe article on Martin Nowak

Panorama interview

Yomiuri article

R.O.M.E.
is currently accepting applications for student researchers

E.T.C.
Evolution and Theology of Cooperation


  • Virtual Labs
  • Charles Darwin
  • Charles Darwin (1809-1882) in his later years. Photo by J. Cameron, 1869.

    Image: © Bettmann/CORBIS

Mission Statement

We are devoted to researching applications of Mathematics and Computer Science to Biology. At the center of a quantitative approach to biology is evolutionary theory as pioneered by Charles Darwin. Concepts of evolutionary biology can be formulated in terms of mathematical equations describing mutation and selection of replicating individuals. We have active research projects on the evolutionary dynamics of infectious agents, cancer cells, altruistic behavior, and human language.

The Program for Evolutionary Dynamics was established in 2003 by the 27th Harvard University President Lawrence Summers following an imaginative proposal by Jeffrey Epstein and Benedict Gross. The center operates under the auspices of Michael Smith, Dean of the Faculty of Arts and Sciences. Martin Nowak, Professor of Mathematics and Biology, is the director.











Nature 20 March 2008 Press coverage:
Winners don't punish

Nature 11 October 2007Press coverage:
Language evolution





















Via Freedom to Coercion: The Emergence of Costly Punishment

Five Rules for the Evolution of Cooperation

A simple rule for the evolution of cooperation on graphs and social networks














Evolution of indirect reciprocity

Dynamics of chronic myeloid leukaemia

Evolutionary dynamics on graphs














Emergence of cooperation and evolutionary stability in finite populations

Evolutionary Dynamics of Biological Games

Prisoners of the dilemma